(1) Free central placentation, synandry, divergent palmately reticulate venation
(2) Mesogamy, fusion of epicarp and thalamus to form rind of the fruit, seed germination epigeal.
(3) Axile placentation, extrafloral nectaries, continuous collenchymatous hypodermal ring in stem.
(4) Bicarpellary gynoecium, compound sieve plate unilocular ovary.
