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*** organs in pteris are produced on

$\begin {array} {1 1} (1)\;Dorsal\: side \: of \: the \: prothallus & \quad (2)\;Ventral\: side\: of \: the \: prothallus \\ (3)\;Margin\: of \: the \: fertile\: leaflets & \quad (4)\;Apices\: of\: sporophytic\: stem \end {array}$

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